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As you might expect, these figures are somewhat subjective and a little hard to determine. However, I can give you one good example that might help.
Microsoft is a large company -- they have about 50,000 users. The average Microsoft employee receives about 200 e-mail messages each day, and sends about 50 e-mail messages each day. Unfortunately, these figures don't take into account attachments or attachment sizes, but at least it gives you a pretty good feel for how many messages you might expect in a large company with active e-mail users.
In terms of financial responsibility, or for that matter legal responsibility, I recommend working with both a risk assessor and a legal advisor. For example, your storage issues may be moot if your legal advisor instructs you to keep e-mail for very short periods of time.
Without knowing a lot more details about your organization, your user base and your service level requirements, it's hard to give you much beyond this general advice.
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